Protoceratidae
Protoceratidae is an extinct family of herbivorous North American artiodactyls that lived during the Eocene through Pliocene. While early members of the group were hornless, in later members males developed elaborate cranial ornamentation. They are variously allied with Ruminantia or Tylopoda.

Nobu Tamura ( http://spinops.blogspot.com ) · CC BY 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- protoceratidPBDB
- Sits inside
- RuminantiamorphaPBDB
- Diet
- browserPBDB
- Life habit
- ground dwellingPBDB
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Artiodactyla
Time the record covers
At least 36 million years on record.
The record covers at least 39.7 Mya to 3.6 Mya. No occurrence read is dated outside 45.9 Mya to 2.58 Mya.
271 occurrences of Protoceratidae on record, most of them in the Eocene. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
271 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
55 formations
- Uinta Formation21
- Sespe Formation19
- Friars Formation · La Jolla Group17
- Chadron Formation17
- Fleming Formation17
- Santiago Formation13
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The named rock units Protoceratidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 31 further formations hold fewer occurrences each and are not listed. 33 occurrences have no formation on record, which is common for older literature.
Protoceratidae and its relatives
EoceneWhere Protoceratidae and its relatives were recorded in the Eocene.
Found in the same rock
2,151 taxa- Brontotheriidae42 sites
- Protoreodon pumilus21 sites
- Equidae20 sites
- Sespedectes singularis19 sites
- Palaeolagus temnodon18 sites
- Camelidae17 sites
- Hesperocyon gregarius17 sites
- Peradectes californicus17 sites
- Trionychidae17 sites
- Aphelops megalodus16 sites
- Ischyromys veterior16 sites
- Leptomeryx speciosus16 sites
- Megalagus brachyodon16 sites
- Rhinocerotidae16 sites
- Testudines16 sites
- Lepisosteus15 sites
- Leptomeryx15 sites
- Pareumys15 sites
- Trigenicus profectus15 sites
- Eomyidae14 sites
- Herpetotherium knighti14 sites
- Leptoreodon14 sites
- Leptoreodon major14 sites
- Leptoreodon stocki14 sites
The Paleobiology Database records 224 sites for Protoceratidae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
101 references- Summary of known occurrences of terrestrial vertebrates from Eocene strata of southern California25
- The Synthetoceratinae (Mammalia, Tylopoda, Protoceratidae)22
- Biostratigraphy of Uintan and Duchesnean land mammal assemblages from the Middle Member of the Sespe Formation, Simi Valley, California14
- Synonymies and reidentifications of North American fossil mammals10
- The Eocene Vertebrates of the Uinta Basin, Utah10
- Eocene-Oligocene faunas of the Cypress Hills Formation, Saskatchewan9
- Mammalian Range Zones in the Chadronian White River Formation at Flagstaff Rim, Wyoming7
- Paleontology and Geology of the Badwater Creek Area, Central Wyoming. Part 14. The Artiodactyls7
- New early Miocene Protoceratids (Mammalia, Artiodactyla) from Panama6
- Miocene and Pliocene Artiodactyls, Texas Gulf Coastal Plain6
- Magnetic stratigraphy and biostratigraphy of the middle Eocene Uinta Formation, Uinta Basin, Utah5
- Magnetostratigraphy and biostratigraphy of the Eocene-Oligocene transition, southwestern Montana5
Each of the 271 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 88 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Protoceratidae
- Source
- Wikipedia/Commons
- Credit
- Nobu Tamura ( http://spinops.blogspot.com )
- Licence
- CC BY 3.0
- Commercial use
- the licence allows it
No 3D scan of Protoceratidae was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itRuminantiamorpha through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Marsh 1891