Lepidocoleus
Lepidocoleus is a genus of extinct armored annelid worm in the class Machaeridia. Two notable species are L. caliburnus or the "Excalibur worm", and L. shurikenus, or the "shuriken worm". The creature had a "suit" of armor running down its body in the form of overlapping calcite crystals. This worm probably lived in shallow water reefs feeding on organic waste. It lived from the Hirnantian of the upper Ordovician to the Famennian of the Devonian.

via iDigBio · CC0 · via iDigBio. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- LepidocoleidaePBDB
- Diet
- grazerPBDB, for Machaeridia
- Life habit
- epifaunalPBDB, for Machaeridia
- Motility
- actively mobilePBDB, for Machaeridia
- Vision
- limitedPBDB, for Polychaeta
Time the record covers
At least 74 million years on record.
The record covers at least 468 Mya to 393.47 Mya. No occurrence read is dated outside 477.1 Mya to 387.95 Mya.
40 occurrences of Lepidocoleus on record, most of them in the Ordovician. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
40 occurrences- Ordovician75%
- Silurian10%
- Devonian15%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
23 formations
- Kope Formation5
- Králuv Dvur Formation4
- Fjacka Shale Formation3
- Waynesville Formation2
- Fairview Formation2
- Utica Shale Formation1
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The named rock units Lepidocoleus has been recovered from, largest first. Drawn where the rock is now, not where it formed. 6 occurrences have no formation on record, which is common for older literature.
Lepidocoleus and its relatives
OrdovicianWhere Lepidocoleus and its relatives were recorded in the Ordovician.
Found in the same rock
548 taxa- Climacograptus12 sites
- Isotelus12 sites
- Ostracoda12 sites
- Ctenodonta11 sites
- Ambonychia10 sites
- Flexicalymene10 sites
- Iocrinus10 sites
- Nuculites10 sites
- Serpulites10 sites
- Arthropoma9 sites
- Cornulites9 sites
- Cryptolithus9 sites
- Modiolopsis9 sites
- Petrocrania9 sites
- Rafinesquina9 sites
- Sinuites9 sites
- Trematis9 sites
- Treptoceras9 sites
- Zygospira9 sites
- Ceramoporella8 sites
- Ceratocephala8 sites
- Corynotrypa8 sites
- Cyclora8 sites
- Escharopora8 sites
The Paleobiology Database records 36 sites for Lepidocoleus, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
22 references- The fossil fauna of the Ordovician in the Cincinnati Region8
- Earliest occurrence of the Hirnantia Fauna in the Prague Basin (Czech Republic)4
- Ecological Change during the early Emsian (Devonian) in the Tafilalt (Morocco),the Origin of the Ammonoidea,and the First African Pyrgocystid Edrioasteroids,Machaerids and Phyllocarids4
- Gradient ecology of a biotic invasion: biofacies of the type Cincinnatian Series (Upper Ordovician), Cincinnati, Ohio region, USA3
- The Tretaspis Series of the Kullatorp Core3
- Swedish Late Ordovician marine benthic assemblages and their bearing on brachiopod biogeography2
- Zlíchovian faunas with early ammonoids from Morocco and their use for the correlation of the eastern Anti-Atlas and the western Dra Valley1
- Machaeridians from the Lower Silurian (Llandowery, Telychian) of Shorpshire, England1
- The Walcott-Rust Quarry: Middle Ordovician Trilobite Konservat-Lagerstätten1
- Caradocian-Gedinnian echinoderm associations of Central and Eastern North America1
- Silurian trilobites from southwestern Alaska1
- The paleoecology of a Late Ordovician shale unit from southwest Ohio and southeastern Indiana1
Each of the 40 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 10 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1

Picture of Lepidocoleus
- Source
- iDigBio
- Credit
- via iDigBio
- Licence
- CC0
- Commercial use
- the licence allows it
No 3D scan of Lepidocoleus was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itLepidocoleidae through timeFormations of the OrdovicianThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Faber 1886