Ithonidae
Ithonidae, commonly called moth lacewings and giant lacewings, is a small family of winged insects of the insect order Neuroptera. The family contains a total of ten living genera, and over a dozen extinct genera described from fossils. The modern Ithonids have a notably disjunct distribution, while the extinct genera had a more global range. The family is considered one of the most primitive living neuropteran families. The family has been expanded twice, first to include the genus Rapisma, formerly placed in the monotypic family Rapismatidae, and then in 2010 to include the genera that had been placed into the family Polystoechotidae. Both Rapismatidae and Polystoechotidae have been shown to nest into Ithonidae sensu lato. The larvae of ithonids are grub-like, subterranean and likely phytophagous.

Kesavamurthy N · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- moth lacewingPBDB
- Sits inside
- IthonoideaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 164 million years on record.
The record covers at least 201.4 Mya to 37 Mya. No occurrence read is dated outside 205.7 Mya to 33.9 Mya.
86 occurrences of Ithonidae on record, most of them in the Eocene. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
86 occurrences- Late Triassic2%
- Early Jurassic15%
- Middle Jurassic5%
- Late Jurassic13%
- Early Cretaceous13%
- Late Cretaceous2%
- Eocene50%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
19 formations
- Fur Formation21
- Klondike Mountain Formation11
- Daohugou Formation10
- Posidonia Shale Formation7
- Tranquille Formation · Kamloops Group4
- Yixian Formation · Jehol Group4
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The named rock units Ithonidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 9 occurrences have no formation on record, which is common for older literature.
Ithonidae and its relatives
EoceneWhere Ithonidae and its relatives were recorded in the Eocene.
Found in the same rock
2,123 taxa- Palaeopsychops latifasciatus8 sites
- Palaeopsychops angustifasciatus7 sites
- Palaeopsychops abruptus6 sites
- Coleoptera5 sites
- Necrotaulius parvulus5 sites
- Panorpidium minima5 sites
- Heterophlebia buckmani4 sites
- Panorpidium geinitzi4 sites
- Panorpidium media4 sites
- Prohemerobius prodromus4 sites
- Rasnitsyneura septemvirgatus4 sites
- Archegocimicidae3 sites
- Archijassus heeri3 sites
- Blattula langfeldti3 sites
- Crusopimpla violina3 sites
- Eoptychoptera simplex3 sites
- Gryllidae3 sites
- Heterothemis brodiei3 sites
- Juraphlebia eugeniae3 sites
- Lasiosmylus newi3 sites
- Locustopsis bernstorffi3 sites
- Mesobittacus minutus3 sites
- Neorthophlebia maculipennis3 sites
- Okanagrion3 sites
The Paleobiology Database records 54 sites for Ithonidae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
35 references- Silky lacewings (Neuroptera: Psychopsidae) from the Eocene-Paleocene transition of Denmark with a review of the fossil record and comments on phylogeny and zoogeography20
- Tertiary Giant Lacewings (Neuroptera: Polystoechotidae): revision and description of new taxa from Western North America and Denmark14
- Die Insektenfauna des Ostniedersachsischen Oberen Lias5
- New Eocene giant lacewings with once-banded wings (Neuroptera: Polystoechotidae) of the Okanagan Highlands at McAbee, Canada, and Republic, United States of America3
- Revision of Epigambriinae Handlirsch, stat. nov., a subfamily of Early Jurassic Ithonidae s.l. (Neuroptera)3
- A new species of Lasiosmylus from the Early Cretaceous, China clarifies its genus-group placement in Ithonidae (Neuroptera)3
- New representatives of the Neuropterans (Neuroptera) from the Jurassic of the Karatau region3
- Two new genera of giant lacewings (Insecta, Neuroptera, Ithonidae) from the Middle Jurassic of China2
- New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae)2
- Lichen mimesis in mid-Mesozoic lacewings2
- A new genus and first Cenozoic fossil record of moth lacewings (Neuroptera: Ithonidae) from the Early Eocene of North America2
- First record of fossil "rapismatid-like" Ithonidae (Insecta, Neuroptera) from the Lower Cretaceous Crato Formation of Brazil2
Each of the 86 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 23 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Ithonidae
- Source
- Wikipedia/Commons
- Credit
- Kesavamurthy N
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
No 3D scan of Ithonidae was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Ithonidae”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itIthonoidea through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Newman 1853