Isoetaceae
Isoetaceae is a family including living quillworts (Isoetes) and comparable extinct herbaceous lycopsids (Tomiostrobus). It was previously thought to have evolved by reduction in size from arborsecent Isoetales such as Pleuromeia, but the discovery of Isoetes beestonii from the Early Triassic shows that the growth habits of modern quillworts predate Pleuromeia and point to the family being a survivor of the Permian–Triassic extinctions.

Retallack · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- IsoetalesPBDB
- Diet
- "photoautotroph"PBDB, for Streptophyta
- Environment
- freshwater,terrestrialPBDB, for Lycopodiophyta
Time the record covers
At least 243 million years on record.
The record covers at least 243.43 Mya to 11.7 ka. No occurrence read is dated outside 244.4 Mya to 0 Mya.
157 occurrences of Isoetaceae on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
157 occurrences- Middle Triassicunder 1%
- Middle Jurassic2%
- Late Jurassic3%
- Early Cretaceous10%
- Late Cretaceous16%
- Paleocene14%
- Eocene3%
- Oligocene1%
- Miocene3%
- Pliocene2%
- Pleistocene45%
- Holoceneunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
27 formations
- Monte Chico Formation13
- Bhuj Formation12
- Cerro Cazador Formation9
- Lefipán Formation7
- Rintoul Creek Formation · Strzelecki Group5
- Golden Valley Formation4
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The named rock units Isoetaceae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 3 further formations hold fewer occurrences each and are not listed. 79 occurrences have no formation on record, which is common for older literature.
Isoetaceae and its relatives
PleistoceneWhere Isoetaceae and its relatives were recorded in the Pleistocene.
Found in the same rock
1,268 taxa- Isoetes76 sites
- Poaceae66 sites
- Chenopodiaceae60 sites
- Ericaceae55 sites
- Quercus47 sites
- Pinus45 sites
- Taraxacum45 sites
- Apiaceae41 sites
- Aster40 sites
- Cupressaceae40 sites
- Plantago40 sites
- Botryococcus39 sites
- Cyperaceae39 sites
- Artemisia38 sites
- Cyathidites minor36 sites
- Brassicaceae35 sites
- Podocarpidites32 sites
- Baculatisporites comaumensis31 sites
- Echinosporis31 sites
- Olea31 sites
- Laevigatosporites ovatus29 sites
- Cyatheacidites annulatus27 sites
- Ephedra26 sites
- Phyllocladidites mawsonii26 sites
The Paleobiology Database records 146 sites for Isoetaceae, and this reads all of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
37 references- Early Pleistocene vegetation change in upland south-eastern Australia21
- Palinología del entorno cretácico/paleógeno del SO de la provincia de Santa Cruz: significado estratigráfico y paleoambiental18
- Site U138911
- Site U138610
- A Catalogue of Fossil Plants from India Part 2: Palaeozoic and Mesozoic Megafossils10
- Site U13858
- Site U13878
- Site U13917
- Paleocene flora of the Rocky Mountains and Great Plains7
- Estudio palinológico de la Formación Cerro Cazador (Cretácico tardío) de la Provincia de Santa Cruz, Argentina6
- Site U13906
- Cretaceous/Paleogene Floral Turnover in Patagonia: Drop in Diversity, Low Extinction, and a Classopollis Spike6
Each of the 157 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 25 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Isoetaceae
- Source
- Wikipedia/Commons
- Credit
- Retallack
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan of Isoetaceae was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Isoetaceae”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itIsoetales through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Dumortier 1829