Homunculus
Homunculus is an extinct genus of New World monkey that lived in Patagonia during the Miocene. Two species are known: Homunculus patagonicus and Homunculus vizcainoi, which are known from material found in the Santa Cruz Formation in the far south of Argentina. Reaching a latitude of ~55°S at the time it lived, it is the southernmost non-human primate ever recorded.

Jonathan M.G. Perry, Sergio F. Vizcaíno, M. Susana Bargo, Néstor Toledo, Kellyn Sanders, Edwin Dickinson, Paul E. Mor… · CC BY 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- HomunculidaePBDB
- Diet
- omnivorePBDB, for Primates
- Life habit
- arborealPBDB, for Primates
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Eutheria
Time the record covers
At least 3.7 million years on record.
The record covers at least 17.5 Mya to 13.8 Mya. No occurrence read is dated outside 21 Mya to 11 Mya.
23 occurrences of Homunculus on record, most of them in the Miocene. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
23 occurrences- Miocene100%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
2 formations
Loading coastlines…
The named rock units Homunculus has been recovered from, largest first. Drawn where the rock is now, not where it formed. 1 occurrence has no formation on record, which is common for older literature.
Homunculus and its relatives
MioceneWhere Homunculus and its relatives were recorded in the Miocene.
Found in the same rock
192 taxa- Cladosictis patagonica10 sites
- Protypotherium praerutilum10 sites
- Sipalocyon gracilis10 sites
- Borhyaena tuberata9 sites
- Hapalops8 sites
- Interatherium extensus8 sites
- Acdestis oweni7 sites
- Adinotherium7 sites
- Neoreomys australis7 sites
- Peltephilus pumilus7 sites
- Acarechimys minutus6 sites
- Adinotherium ovinum6 sites
- Nesodon imbricatus6 sites
- Protypotherium australe6 sites
- Prozaedyus proximus6 sites
- Thoatherium minusculum6 sites
- Acarechimys minutissimus5 sites
- Astrapotherium5 sites
- Astrapotherium magnum5 sites
- Eucholoeops ingens5 sites
- Microbiotherium tehuelchum5 sites
- Nesodon5 sites
- Palaeothentes lemoinei5 sites
- Palaeothentes minutus5 sites
The Paleobiology Database records 17 sites for Homunculus, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
11 references- Paleobiology of Santacrucian primates10
- New Primates From the Río Santa Cruz and Río Bote (Early-Middle Miocene), Santa Cruz Province, Argentina4
- Previously unknown fossil platyrrhines (Primates) of Patagonia from the Tournouër collection at the Muséum national d'Histoire naturelle, Paris1
- Fossil localities of the Santa Cruz Formation (Early Miocene, Patagonia, Argentina) prospected by Carlos Ameghino in 1887 revisited and the location of the Notohippidian1
- Oldest known cranium of a juvenile New World monkey (Early Miocene, Patagonia, Argentina): Implications for the taxonomy and the molar eruption pattern of early platyrrhines1
- A review of the paleoenvironment and paleoecology of the Miocene Santa Cruz Formation1
- Two new fossil vertebrate localities in the Santa Cruz Formation (late early – early middle Miocene, Argentina), ~ 51° South latitude1
- New primate genus from the Miocene of Argentina1
- Bioestratigrafía de la Formación Santa Cruz (Mioceno Inferior) en el extremo sudeste de la Patagonia1
- Comparative Anatomy of Platyrrhine Mandibular Cheek Teeth dpm4, pm4, m1 with Particular Reference to Those of Homunculus (Cebidae), and Comments on Platyrrhine Origins1
- Ceboid monkeys from the Miocene of Colombia1
Each of the 23 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content3
Picture of Homunculus
- Source
- Wikipedia/Commons
- Credit
- Jonathan M.G. Perry, Sergio F. Vizcaíno, M. Susana Bargo, Néstor Toledo, Kellyn Sanders, Edwin Dickinson, Paul E. Mor…
- Licence
- CC BY 4.0
- Commercial use
- the licence allows it
CT scan
- Source
- MorphoSource
- Licence
- In Copyright
- Commercial use
- cannot be read from the licence given — check the record
- Specimen
- Homunculus patagonicus · mpm:pv:19426
3D model
- Source
- Sketchfab
- Credit
- The WarVet
- Licence
- CC Attribution
- Commercial use
- cannot be read from the licence given — check the record
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itHomunculidae through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Ameghino 1891