Grylloblattodea
Notoptera, also known as Xenonomia, is an order of insects belonging to Polyneoptera. It contains two living suborders, Mantophasmatidae (gladiators) native to Southern Africa, and Grylloblattidae native to cold montane environments in the Northern Hemisphere. Both are wingless.

P.E. Bragg · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- ice crawlerPBDB
- Sits inside
- NotopteraPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 216 million years on record.
The record covers at least 316.9 Mya to 100.5 Mya. No occurrence read is dated outside 318.7 Mya to 93.9 Mya.
555 occurrences of Grylloblattodea on record, most of them in the Permian. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
555 occurrences- Carboniferous5%
- Permian75%
- Early Triassic1%
- Middle Triassic10%
- Late Triassicunder 1%
- Early Jurassic3%
- Middle Jurassicunder 1%
- Late Jurassic3%
- Late Cretaceous1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
60 formations
- Wellington Formation · Cimarron Group134
- Koshelevka Formation62
- Madygen Formation45
- Mitina Formation45
- Letovice Formation26
- Iva-Gora Beds Formation16
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The named rock units Grylloblattodea has been recovered from, largest first. Drawn where the rock is now, not where it formed. 36 further formations hold fewer occurrences each and are not listed. 27 occurrences have no formation on record, which is common for older literature.
Grylloblattodea and its relatives
PermianWhere Grylloblattodea and its relatives were recorded in the Permian.
Found in the same rock
5,731 taxa- Artinska clara10 sites
- Lisca minuta10 sites
- Lemmatophora typa9 sites
- Artinska ovata8 sites
- Dichentomum tinctum7 sites
- Liomopteridae7 sites
- Permopsylla americana7 sites
- Asthenohymen dunbari6 sites
- Coleoptera6 sites
- Delopsocus elongatus6 sites
- Delopsocus latus6 sites
- Delopterum minutum6 sites
- Heterophlebia buckmani6 sites
- Lecorium elongatum6 sites
- Liomopterum ornatum6 sites
- Paleuthygramma acuta6 sites
- Panorpidium geinitzi6 sites
- Panorpidium minima6 sites
- Parablattelytron rectum6 sites
- Permopsocus latipennis6 sites
- Phenopterum elongatum6 sites
- Protohymen permianus6 sites
- Tillyardembia antennaeplana6 sites
- Aktassoblatta fusca5 sites
The Paleobiology Database records 166 sites for Grylloblattodea, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
134 references- Paleozoiskie nasekomye kuznetskogo basseina50
- Types of Permian Insects, Part III: Megasecoptera, Orycloblattinidae and Protorthoptera34
- The Fauna of Grylloblattid Insects (Grylloblattida) of the Lower Permian Locality of Tshekarda29
- Permian insects of Moravia. Part II - Liomopteridea25
- Neue Untersuchungen über die fossilen Insekten mit Ergänzungen und Nachträgen sowie Ausblicken auf phylogenetische, palaeogeographische und allgemein biologische Probleme. I. Teil20
- New Triassic grylloblattids from Kirghizia (Insecta, Grylloblattida)17
- The Lower Permian insects of Kansas, Part 7: the Order Protoperlaria17
- The Lower Permian insects of Kansas. Part 10. The order Protorthoptera: the family Liomopteridae and its relatives15
- New Mesozoic Grylloblattid insects (Grylloblattida) from Central Asia14
- New grylloblattid insects (Insecta: Grylloblattida) from the Gres a Voltzia of the Vosges (Middle Triassic of France)11
- The Lower Permian Insects of Kansas. Part 8: Additional Megasecoptera, Protodonata, Odonata, Homoptera, Psocoptera, Protelytroptera, Plectoptera and Protoperlaria11
- Diversity and variability of grylloblattidan insects (Grylloblattida) from the early Permian Meisenheim Formation of the Saar‑Nahe Basin (SW‑Germany)10
Each of the 555 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 122 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content2
Picture of Grylloblattodea
- Source
- Wikipedia/Commons
- Credit
- P.E. Bragg
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Notoptera”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itNotoptera through timeFormations of the PermianThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Walker 1914)