Engraulis
Engraulis is a genus of anchovies, currently containing 9 species. Species in this genus are found in the Pacific and Atlantic Oceans, as well as the Mediterranean Sea, and several of them are commercially important.

Uncredited · Public Domain · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- EngraulidaePBDB
- Diet
- carnivorePBDB, for Clupeiformes
- Life habit
- nektonicPBDB, for Clupeiformes
- Motility
- actively mobilePBDB, for Clupeiformes
Time the record covers
At least 33 million years on record.
The record covers at least 33.9 Mya to 11.7 ka. No occurrence read is dated outside 37.71 Mya to 0 Mya.
38 occurrences of Engraulis on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
38 occurrences- Eocene3%
- Miocene8%
- Pliocene28%
- Pleistocene59%
- Holocene3%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
9 formations
- Palos Verdes Sand Formation4
- Lindos Bay Formation3
- Gutingkeng Formation3
- Yokosuka Formation2
- Otuma Formation1
- Gatun Formation1
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The named rock units Engraulis has been recovered from, largest first. Drawn where the rock is now, not where it formed. 21 occurrences have no formation on record, which is common for older literature.
Engraulis and its relatives
PleistoceneWhere Engraulis and its relatives were recorded in the Pleistocene.
Found in the same rock
844 taxa- Maurolicus muelleri10 sites
- Cymatogaster aggregata9 sites
- Galeorhinus galeus9 sites
- Genyonemus lineatus9 sites
- Gobius9 sites
- Porichthys notatus9 sites
- Seriphus politus9 sites
- Trachurus9 sites
- Ceratoscopelus maderensis8 sites
- Conger8 sites
- Diaphus8 sites
- Micromesistius poutassou8 sites
- Raja8 sites
- Triakis semifasciata8 sites
- Citharichthys sordidus7 sites
- Citharichthys stigmaeus7 sites
- Myliobatis californica7 sites
- Squatina californica7 sites
- Bregmaceros albyi6 sites
- Carcharodon carcharias6 sites
- Chilara taylori6 sites
- Citharichthys6 sites
- Cottidae6 sites
- Electrona rissoi6 sites
The Paleobiology Database records 36 sites for Engraulis, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
20 references- 541. Some Pleistocene fish-otoliths from the Boso and Miura peninsulas6
- Données récentes sur Les téléostéens du Miocène et du Pliocéne d'Italie4
- Early Pliocene otolith assemblages from the outer-shelf environment reveal the establishment of mesopelagic fish fauna over 3 million years ago in southwestern Taiwan3
- Pleistocene marine fish invasions and paleoenvironmental reconstructions in the eastern Mediterranean3
- Biogeography and climatology of Pliocene and Messinian fossil fish of eastern-central Italy3
- Fish remains, mostly otoliths and teeth, from the Palos Verdes Sand (Late Pleistocene) of California3
- Fish otolith record reveals possible tropical-subtropical fish community in temperate Japan during the exceptionally warm Last Interglacial period2
- Ricerche sui Pesci Fossili di Chiavòn (Strati di Sotzka -- Miocene Inferiore)2
- The Zanclean marine fish fauna and palaeoenvironmental reconstruction of a coastal marine setting in the eastern Mediterranean1
- Reconstruction of the fossil marine bony fish fauna (Teleostei) from the Eocene to Pleistocene of New Zealand by means of otoliths. With studies of Recent congroid, morid and trachinoid otoliths1
- Fossil evidence for evolution of the shape and color of penguin feathers1
- Bathymetric distribution of Miocene to Pleistocene Caribbean teleostean fishes from the coast of Panama and Costa Rica1
Each of the 38 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 8 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content2
Picture of Engraulis
- Source
- Wikipedia/Commons
- Credit
- nobody named on the record
- Licence
- Public Domain
- Commercial use
- the licence allows it
CT scan
- Source
- MorphoSource
- Credit
- Bell Museum fishes
- Licence
- In Copyright
- Commercial use
- cannot be read from the licence given — check the record
- Specimen
- Engraulis japonicus · JFBM:Fishes:48201
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Engraulis”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itEngraulidae through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Cuvier 1817