Empididae
Empididae is a family of flies with over 3,000 described species occurring worldwide in all the biogeographic realms but the majority are found in the Holarctic. They are mainly predatory flies like most of their relatives in the Empidoidea, and exhibit a wide range of forms but are generally small to medium-sized, non-metallic and rather bristly.

André Karwath aka Aka · CC BY-SA 2.5 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- dagger flyPBDB
- Sits inside
- EmpidoideaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 165 million years on record.
The record covers at least 165.3 Mya to 11.7 ka. No occurrence read is dated outside 170.9 Mya to 0 Mya.
112 occurrences of Empididae on record, most of them in the Eocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
112 occurrences- Middle Jurassicunder 1%
- Late Jurassic5%
- Early Cretaceous4%
- Late Cretaceous4%
- Eocene70%
- Oligocene10%
- Miocene4%
- Holoceneunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
14 formations
- Florissant Formation18
- Karabastau Formation6
- Rott Formation6
- Yixian Formation · Jehol Group2
- Niveau du gypse d'Aix Formation2
- Raritan Formation · Potomac Group2
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The named rock units Empididae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 68 occurrences have no formation on record, which is common for older literature.
Empididae and its relatives
EoceneWhere Empididae and its relatives were recorded in the Eocene.
Found in the same rock
3,880 taxa- Formicidae6 sites
- Brachycera5 sites
- Coleoptera5 sites
- Dolichopodidae5 sites
- Aphaenogaster mayri4 sites
- Codemus synaptoides4 sites
- Corynoptera klebsii4 sites
- Ctenobethylus goepperti4 sites
- Dicranomyia (Dicranomyia) lobata4 sites
- Distenorrhinus (Distenorrhinus) pallidirostris4 sites
- Entimini4 sites
- Eoptychopterina karatavica4 sites
- Gesomyrmex hoernesi4 sites
- Hemiptera4 sites
- Kararhynchus occiduus4 sites
- Lasius schiefferdeckeri4 sites
- Liassothrips crassipes4 sites
- Liometopum miocenicum4 sites
- Macrochile spectrum4 sites
- Miomyrmex impactus4 sites
- Nylanderia pygmaea4 sites
- Pachycondyla succinea4 sites
- Paramesopsocus adibi4 sites
- Plagiolepis klinsmanni4 sites
The Paleobiology Database records 44 sites for Empididae, and this reads all of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
41 references- Monographie des Empidae de l'ambre de la Baltique et catalogue bibliographique complet sur les diptères fossiles de cette résine38
- A Report on Some Miocene Diptera from Florissant, Colorado12
- A brief review of brachycerous flies (Diptera, Brachycera) in the Mesozoic, with descriptions of some curious taxa6
- Ueber den Bernstein und die Bernsteinfauna6
- Neue Dipteren (Brachycera et Cyclorhapha) aus dem Oberoligozän von Rott5
- Brachyceran Diptera in Cretaceous ambers and Mesozoic diversification of the Eremoneura3
- Beschreibung einer Art der Gattung Heleodromia (Diptera, Empididae, Hemerodromiinae) aus dem Bitterfelder Bernstein3
- Die Insecten und Spinnen der Vorwelt mit steter Berücksichtigung der lebenden Insekten und Spinnen3
- Nota sobre algunos insectos fósiles de Ribesalbes (Castellón)2
- Miocene fossil insects2
- Révision des diptères fossiles types de Loew conservés au Musée Provincial de Koenigsberg2
- Proclinopyga ulrichi sp. nov.: the first fossil aquatic dance fly of the subfamily Clinocerinae (Diptera: Empididae)1
Each of the 112 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 29 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Empididae
- Source
- Wikipedia/Commons
- Credit
- André Karwath aka Aka
- Licence
- CC BY-SA 2.5
- Commercial use
- the licence allows it
No 3D scan of Empididae was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Empididae”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itEmpidoidea through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Latreille 1804