Borophagus littoralis
Borophagus littoralis is an extinct species of the genus Borophagus of the subfamily Borophaginae, a group of canids endemic to North America from the Late Miocene from 11.6 to 9.4 Mya. Borophagus littoralis existed for approximately 2.2 million years.

Patricia Holroyd · CC BY 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- BorophagusPBDB
- Diet
- carnivorePBDB, for Borophagus
- Life habit
- ground dwellingPBDB, for Canidae
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Eutheria
Time the record covers
At most 7.2 million years on record.
Every occurrence read is dated to somewhere between 12.5 Mya and 5.333 Mya, and nothing in the dating separates them, so that window is what the record shows rather than a span.
8 occurrences of Borophagus littoralis on record, most of them in the Miocene. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
8 occurrences- Miocene100%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
4 formations
Loading coastlines…
The named rock units Borophagus littoralis has been recovered from, largest first. Drawn where the rock is now, not where it formed. 2 occurrences have no formation on record, which is common for older literature.
Borophagus littoralis and its relatives
MioceneWhere Borophagus littoralis and its relatives were recorded in the Miocene.
Found in the same rock
58 taxa- Dinohippus leardi5 sites
- Hipparion forcei4 sites
- Caninae3 sites
- Cupidinimus3 sites
- Leptocyon vafer3 sites
- Perognathus minutus3 sites
- Plioceros3 sites
- Sorex3 sites
- Ustatochoerus californicus3 sites
- Barbourofelis whitfordi2 sites
- Copemys dentalis2 sites
- Cormohipparion occidentale2 sites
- Epicyon haydeni2 sites
- Martinogale2 sites
- Paronychomys2 sites
- Pliohippus tehonensis2 sites
- Procyonidae2 sites
- Scapanus shultzi2 sites
- Serbelodon burnhami2 sites
- Aelurodon taxoides1 site
- Aepycamelus1 site
- Alilepus hibbardi1 site
- Alluvisorex chasseae1 site
- Barbourofelis loveorum1 site
The Paleobiology Database records 8 sites for Borophagus littoralis, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
5 references- Synonymies and reidentifications of North American fossil mammals2
- Phylogenetic systematics of the Borophaginae (Carnivora: Canidae)2
- Miocene biostratigraphy and biochronology of the Dove Spring Formation, Mojave Desert, California, and characterization of the Clarendonian mammal age (late Miocene) in California2
- Lower Pliocene horses from Black Hawk Ranch, Mount Diablo, California1
- Borophagus littoralis From the Marine Tertiary of California1
Each of the 8 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Borophagus littoralis
- Source
- Wikipedia/Commons
- Credit
- Patricia Holroyd
- Licence
- CC BY 3.0
- Commercial use
- the licence allows it
No 3D scan of Borophagus littoralis was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itBorophagus through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (VanderHoof 1931)