Borophaginae
The extinct Borophaginae form one of three subfamilies found within the canid family. The other two canid subfamilies are the extinct Hesperocyoninae and extant Caninae. Borophaginae, called "bone-crushing dogs", were endemic to North America during the Oligocene to Pliocene and lived roughly 34–2.5 million years ago, existing for about 31.5 million years.

Jeff Kubina from Columbia, Maryland · CC BY-SA 2.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- bone-crushing dogPBDB
- Sits inside
- CanidaePBDB
- Diet
- carnivore, omnivorePBDB, for Canidae
- Life habit
- ground dwellingPBDB, for Canidae
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Eutheria
Time the record covers
At least 29 million years on record.
The record covers at least 31.8 Mya to 2.58 Mya. No occurrence read is dated outside 41.03 Mya to 1.4 Mya.
1,137 occurrences of Borophaginae on record, most of them in the Miocene. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
1,137 occurrences- Eoceneunder 1%
- Oligocene8%
- Miocene86%
- Pliocene5%
- Pleistocene1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
125 formations
- Ash Hollow Formation · Ogallala Group132
- Valentine Formation · Ogallala Group111
- Runningwater Formation · Hemingford Group60
- Barstow Formation55
- Olcott Formation43
- Sharps Formation26
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The named rock units Borophaginae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 101 further formations hold fewer occurrences each and are not listed. 235 occurrences have no formation on record, which is common for older literature.
Borophaginae and its relatives
MioceneWhere Borophaginae and its relatives were recorded in the Miocene.
Found in the same rock
954 taxa- Cynarctoides acridens35 sites
- Desmocyon thomsoni29 sites
- Aphelops megalodus27 sites
- Archaeocyon leptodus27 sites
- Merychyus relictus18 sites
- Metatomarctus canavus18 sites
- Nanotragulus loomisi18 sites
- Teleoceras medicornutum18 sites
- Camelidae17 sites
- Protomarctus optatus16 sites
- Tomarctus hippophaga16 sites
- Leptauchenia major14 sites
- Merychyus minimus14 sites
- Aelurodon taxoides13 sites
- Equidae13 sites
- Leptauchenia decora13 sites
- Leptocyon leidyi13 sites
- Palaeocastor13 sites
- Capacikala gradatus12 sites
- Epicyon saevus12 sites
- Palaeolagus philoi12 sites
- Stenomylus gracilis12 sites
- Ticholeptus zygomaticus12 sites
- Menoceras barbouri11 sites
The Paleobiology Database records 757 sites for Borophaginae, and this reads 240 of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
160 references- Phylogenetic systematics of the Borophaginae (Carnivora: Canidae)613
- Synonymies and reidentifications of North American fossil mammals173
- Vertebrate biostratigraphy of the Ogallala Group in Nebraska27
- Miocene mammals of the Split Rock area, Granite Mountains basin, central Wyoming12
- Review of the Miocene Wounded Knee faunas of southwestern South Dakota12
- Revised chronostratigraphy and biostratigraphy of the John Day Formation (Turtle Cove and Kimberly Members), Oregon, with implications for updated calibration of the Arikareean North American land mammal age11
- Evolutionary trends in the late Miocene hyena-like dog Epicyon (Carnivora, Canidae)10
- The Wood Mountain fauna: an upper Miocene mammalian assemblage from southern Saskatchewan10
- Stratigraphy and biostratigraphy of late Cenozoic deposits in central Sioux County, western Nebraska8
- Miocene biostratigraphy and biochronology of the Dove Spring Formation, Mojave Desert, California, and characterization of the Clarendonian mammal age (late Miocene) in California7
- Stop 14: The Clarendonian faunas of the Texas and Oklahoma panhandles7
- The geology and paleontology of Ashfall Fossil Beds, a late Miocene (Clarendonian) mass-death assemblage, Antelope County and adjacent Knox County, Nebraska, USA6
Each of the 1,137 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 148 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Borophaginae
- Source
- Wikipedia/Commons
- Credit
- Jeff Kubina from Columbia, Maryland
- Licence
- CC BY-SA 2.0
- Commercial use
- the licence allows it
No 3D scan of Borophaginae was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itCanidae through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Simpson 1945