Ballusia
Ballusia was a genus of small bear from the Early Miocene epoch, about 20.5-18 million years ago. Fossil remains attributed to the genus have been uncovered in Europe (Poland) and Asia. The genus Ballusia was established in 1998 on the basis of different fossils originally classified as various species of the genera Ursavus and Hemicyon, with B. elmensis as the type species. The exact relationship of Ballusia to "true" bears are not yet fully understood: many palaeontologists have classified it as a primitive member of Ursinae, but its known skeletal elements have some features in common with the extinct bear subfamily Hemicyoninae. Because of this, some researchers refer Ballusia as "Ursidae incertae sedis". Ginsburg and Morales regarded B. elmenensis as ancestral to Ursavus, as did Marciszak and Lipecki, even though the temporal range of the two genera seems to have overlapped.

Conty · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- UrsinaePBDB
- Diet
- herbivore, carnivorePBDB, for Ursidae
- Life habit
- ground dwellingPBDB, for Ursidae
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Eutheria
Time the record covers
At least 920 thousand years on record.
The record covers at least 16.9 Mya to 15.98 Mya. No occurrence read is dated outside 23.04 Mya to 5.333 Mya.
6 occurrences of Ballusia on record, most of them in the Miocene. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
6 occurrences- Miocene100%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
1 formation
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The named rock units Ballusia has been recovered from, largest first. Drawn where the rock is now, not where it formed. 5 occurrences have no formation on record, which is common for older literature.
Ballusia and its relatives
MioceneWhere Ballusia and its relatives were recorded in the Miocene.
Found in the same rock
236 taxa- Hemicyon5 sites
- Lagopsis4 sites
- Amphictis3 sites
- Anchitherium aurelianense3 sites
- Andegameryx3 sites
- Aureliachoerus aurelianensis3 sites
- Brachyodus3 sites
- Cynelos helbingi3 sites
- Martes3 sites
- Phoberocyon3 sites
- Plithocyon3 sites
- Protaceratherium minutum3 sites
- Pseudaelurus3 sites
- Pseudocyon sansaniensis3 sites
- Semigenetta3 sites
- Steneofiber depereti3 sites
- Styriofelis turnauensis3 sites
- Xenohyus venitor3 sites
- Amphicyon2 sites
- Amphicyon lathanicus2 sites
- Cainotherium lintillae2 sites
- Chainodus intercedens2 sites
- Dorcatherium guntianum2 sites
- Haplocyonoides mordax2 sites
The Paleobiology Database records 6 sites for Ballusia, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
6 references- Le site paléontologique du Grand Morier (Pont-Boutard, Indre-et-Loire, France): contexte géologique et détail biostratigraphique des formations cénozoïques à partir des assemblages de vertébrés fossiles1
- Les fossiles de mammifères miocènes de la colletion de J.-P. Hartmann conservés dans le Musée du Savignéen1
- Die erst Wirbeltierfauna aus der miozanen Braunkohle der Niederrheinischen Bucht (Ville-Schichten, Tagebau Hambach) [The first vertebrate fauna from the Miocene Ville Series of the Lower Rhine Embayment (Hambach open cast mine, western Germany)]1
- La succession des faunes de mammiferes miocenes de Pontigne (Maine-et-Loire, France)1
- Les mammifères de la faune sidérolithique de Vieux-Collonges1
- Die Raubtiere aus dem Mittel-Miocan (Burgidalium) von Wintershof-West bei Eichstatt in Bayern.1
Each of the 6 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Ballusia
- Source
- Wikipedia/Commons
- Credit
- Conty
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan of Ballusia was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Ballusia”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itUrsinae through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Ginsburg and Morales 1998