Therocephalia
Therocephalia is an extinct group of therapsids from the Permian and Triassic periods. The therocephalians ("beast-heads") are named after their large skulls, which, along with the structure of their teeth, suggest that they were carnivores. Like other non-mammalian synapsids, therocephalians were once described as "mammal-like reptiles". Therocephalia is the group most closely related to the cynodonts, which gave rise to the mammals. Indeed, it had been proposed that therocephalians themselves may have given rise to the cynodonts, and therefore that therocephalians as recognised are paraphyletic in relation to cynodonts and so not a clade. Conventionally, however, Therocephalia is regarded as the sister clade of Cynodontia, together forming the clade Eutheriodontia.

w:ru:Участник:ДиБгд (dmitrchel@mail.ru) · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- TherapsidaPBDB
- Diet
- carnivorePBDB
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Amniota
Time the record covers
At least 20 million years on record.
The record covers at least 264.28 Mya to 243.43 Mya. No occurrence read is dated outside 274.4 Mya to 237 Mya.
344 occurrences of Therocephalia on record, most of them in the Permian. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
344 occurrences- Permian78%
- Early Triassic13%
- Middle Triassic10%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
28 formations
- Balfour Formation · Beaufort Group33
- Abrahamskraal Formation · Beaufort Group28
- Burgersdorp Formation · Beaufort Group21
- Teekloof Formation · Beaufort Group11
- Urpalov Formation · Kotelnich Group9
- Normandien Formation · Beaufort Group8
Loading coastlines…
The named rock units Therocephalia has been recovered from, largest first. Drawn where the rock is now, not where it formed. 4 further formations hold fewer occurrences each and are not listed. 178 occurrences have no formation on record, which is common for older literature.
Therocephalia and its relatives
PermianWhere Therocephalia and its relatives were recorded in the Permian.
Found in the same rock
404 taxa- Theriognathus microps19 sites
- Moschorhinus kitchingi18 sites
- Lystrosaurus declivis9 sites
- Bauria cynops8 sites
- Lystrosaurus murrayi8 sites
- Lystrosaurus curvatus7 sites
- Procynosuchus delaharpeae7 sites
- Microgomphodon oligocynus6 sites
- Prolacerta broomi6 sites
- Dicynodon5 sites
- Isadia aristoviensis5 sites
- Procolophon trigoniceps5 sites
- Tetracynodon darti5 sites
- Thrinaxodon liorhinus5 sites
- Anteosaurus magnificus4 sites
- Diictodon4 sites
- Diictodon feliceps4 sites
- Emydops4 sites
- Glanosuchus macrops4 sites
- Kannemeyeria simocephala4 sites
- Karpinskiosaurus secundus4 sites
- Lystrosaurus4 sites
- Lystrosaurus maccaigi4 sites
- Olivierosuchus parringtoni4 sites
The Paleobiology Database records 130 sites for Therocephalia, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
75 references- The distribution of the Karroo vertebrate fauna109
- The fauna of the Tapinocephalus Zone (Beaufort Beds of the Karoo)31
- Permian and Triassic therocephals (Eutherapsida) of Eastern Europe29
- Revision of the first therocephalian, Theriognathus Owen (Therapsida: Whaitsiidae), and implications for cranial ontogeny and allometry in nonmammaliaform eutheriodonts22
- On the Whatsiidae, a family of therocephalian mammal-like reptiles22
- Body size and growth patterns in the therocephalian Moschorhinus kitchingi (Therapsida: Eutheriodontia) before and after the end-Permian extinction in South Africa16
- New Material of Microgomphodon oligocynus (Eutherapsida, Therocephalia) and the Taxonomy of Southern African Bauriidae8
- On Bauria cynops Broom6
- Mammal-like reptiles from Russia5
- New whaitsioids (Therapsida: Therocephalia) from the Teekloof Formation of South Africa and therocephalian diversity during the end Guadalupian extinction4
- Reassessment of the morphology and paleobiology of the therocephalian Tetracynodon darti (Therapsida), and the phylogenetic relationships of Baurioidea4
- A new skeleton of the therocephalian synapsid Olivierosuchus parringtoni from the Lower Triassic South African Karoo Basin4
Each of the 344 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 63 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Therocephalia
- Source
- Wikipedia/Commons
- Credit
- w:ru:Участник:ДиБгд (dmitrchel@mail.ru)
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
No 3D scan of Therocephalia was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itTherapsida through timeFormations of the PermianThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Broom 1903)