Pipunculidae
Pipunculidae is a family of flies (Diptera) commonly termed big-headed flies, a reference to the large (holoptic) eyes, which cover nearly the entire head. The family is found worldwide and more than 1300 species have been described.

This image is created by user Dick Belgers at Waarneming.nl , a source of nature observations in the Netherlands. · CC BY 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- big-headed flyPBDB
- Sits inside
- CyclorrhaphaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 27 million years on record.
The record covers at least 48.07 Mya to 20.45 Mya. No occurrence read is dated outside 56 Mya to 13.82 Mya.
15 occurrences of Pipunculidae on record, most of them in the Eocene. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
15 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
3 formations
- Tranquille Formation · Kamloops Group2
- Florissant Formation1
- Klondike Mountain Formation1
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The named rock units Pipunculidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 11 occurrences have no formation on record, which is common for older literature.
Pipunculidae and its relatives
EoceneWhere Pipunculidae and its relatives were recorded in the Eocene.
Found in the same rock
2,354 taxa- Arctogephyra agilis4 sites
- Agraylea spathifera3 sites
- Aphaenogaster sommerfeldti3 sites
- Carebara antiqua3 sites
- Corynoptera klebsii3 sites
- Dicranomyia (Dicranomyia) lobata3 sites
- Elephantomyia (Elephantomyia) longirostris3 sites
- Eotemnothorax gracilis3 sites
- Gesomyrmex hoernesi3 sites
- Harpactosphecion gracile3 sites
- Kroeberiella pinguis3 sites
- Lasius schiefferdeckeri3 sites
- Lype sericea3 sites
- Macrochile spectrum3 sites
- Nylanderia pygmaea3 sites
- Palaeohelicopsyche serricornis3 sites
- Palaeopoecilostola longicornis3 sites
- Palaeopoecilostola speciosa3 sites
- Plagiolepis kuenowi3 sites
- Plectrocnemia lata3 sites
- Prenolepis henschei3 sites
- Procerapachys annosus3 sites
- Prosphyracephala succini3 sites
- Sylvicola thiriona3 sites
The Paleobiology Database records 7 sites for Pipunculidae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
7 references- The fossil Pipunculidae4
- Les Pipunculidae de l'ambre3
- Early Eocene big headed flies (Diptera: Pipunculidae) from the Okanagan Highlands, western North America2
- Micro-CT studies of amber inclusions reveal internal genitalic features of big-headed flies, enabling a systematic placement of Metanephrocerus Aczél, 1948 (Insecta: Diptera: Pipunculidae)2
- Short note on fossil Pipunculidae (Diptera) from Dominican amber2
- Early Eocene insects from Quilchena, British Columbia, and their paleoclimatic implications1
- Grundlagen einer Monographie der Dorilaiden (Diptera). Dorilaiden-Studien VI1
Each of the 15 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Pipunculidae
- Source
- Wikipedia/Commons
- Credit
- This image is created by user Dick Belgers at Waarneming.nl , a source of nature observations in the Netherlands.
- Licence
- CC BY 3.0
- Commercial use
- the licence allows it
No 3D scan of Pipunculidae was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itCyclorrhapha through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Walker 1834