Pinaceae
The Pinaceae, or pine family, is a large family of conifers, including many of the well-known conifers of commercial importance such as cedars, firs, hemlocks, larches, pines, and spruces. The family is included in the order Pinales, formerly known as Coniferales. Pinaceae have distinctive cones with woody scales bearing typically two ovules, and are supported as monophyletic by both morphological trait and genetic analysis. They are the largest extant conifer family in species diversity, with between 220 and 250 species in 11 genera, and the second-largest in geographical range, found in most of the Northern Hemisphere, with the majority of the species in temperate climates, but ranging from subarctic to tropical. The family often forms the dominant component of boreal, coastal, and montane forests. One species, Pinus merkusii, grows just south of the equator in Southeast Asia. Major centres of diversity are found in the mountains of southwest China, Mexico, central Japan, and California.

Marshmallow from Seattle, WA, USA · CC BY 2.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- PinalesPBDB
- Diet
- "photoautotroph"PBDB, for Streptophyta
Time the record covers
At least 306 million years on record.
The record covers at least 307 Mya to 11.7 ka. No occurrence read is dated outside 358.86 Mya to 0 Mya.
1,736 occurrences of Pinaceae on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
1,736 occurrences- Carboniferous2%
- Permian2%
- Early Triassicunder 1%
- Middle Triassicunder 1%
- Late Triassicunder 1%
- Early Jurassicunder 1%
- Middle Jurassicunder 1%
- Late Jurassicunder 1%
- Early Cretaceous3%
- Late Cretaceous3%
- Paleocene3%
- Eocene12%
- Oligocene7%
- Miocene27%
- Pliocene7%
- Pleistocene28%
- Holocene3%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
173 formations
- Chloropagus Formation59
- Fort Union Formation31
- Yaw Formation28
- Hell Creek Formation20
- Middlegate Formation18
- Turtel Mountain Formation16
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The named rock units Pinaceae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 149 further formations hold fewer occurrences each and are not listed. 1,115 occurrences have no formation on record, which is common for older literature.
Pinaceae and its relatives
PleistoceneWhere Pinaceae and its relatives were recorded in the Pleistocene.
Found in the same rock
2,900 taxa- Laevigatosporites67 sites
- Pinus59 sites
- Deltoidospora58 sites
- Pinuspollenites52 sites
- Microreticulatisporites51 sites
- Leiotriletes50 sites
- Taxodium50 sites
- Cicatricosisporites dorogensis48 sites
- Coniferales48 sites
- Pediastrum48 sites
- Caryapollenites47 sites
- Converrucosisporites47 sites
- Triporopollenites47 sites
- Pityosporites46 sites
- Thompsonipollis magnificus46 sites
- Casurinidites pulcher45 sites
- Liliacidites45 sites
- Picea42 sites
- Gleicheniidites41 sites
- Tricolpopollenites41 sites
- Ovoidites40 sites
- Margocolporites39 sites
- Momipites39 sites
- Nudopollis terminalis39 sites
The Paleobiology Database records 771 sites for Pinaceae, and this reads 240 of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
235 references- A Catalogue of Fossil Plants from India Part 6: Cenozoic (Quaternary) Palynology and Palaeobotany121
- Die mittelmiozaenen und obermiozaenen Floren aus der Meuroer und Raunoer Folge in der Lausitz. III. Fundstellen und Palaeobiologie117
- Site U1344108
- Site U134374
- Distribution of pollen and spores in a subsurface marine Wilcox (Paleocene-Eocene) section in southwest Louisiana63
- The Miocene Purple Mountain Flora of Western Nevada56
- Biostratigraphy of selected K/T boundary sections in southwestern North Dakota, USA: toward a refinement of palynological identification criteria48
- Die oberoligozanen Floren am Nordrand der Sachsischen Lausitz43
- A catalogue of Cenozoic megafossil plants in China41
- Site U134535
- Tretichnye Flory Zapadnoi Sibiri33
- Niche evolution versus niche conservatism and habitat loss determine persistence and extirpation in late Neogene European Fagaceae32
Each of the 1,736 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 223 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Pinaceae
- Source
- Wikipedia/Commons
- Credit
- Marshmallow from Seattle, WA, USA
- Licence
- CC BY 2.0
- Commercial use
- the licence allows it
No 3D scan of Pinaceae was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Pinaceae”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itPinales through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Rudolphi 1830