Ommatidae
The Ommatidae are a family of beetles in the suborder Archostemata. The Ommatidae are considered the extant beetle family that has most ancestral characteristics. There are only seven extant species, confined to Australia and South America. However, the geographical distribution was much wider during the Mesozoic spanning across Eurasia, South America, and Australia, suggesting that they were widespread on Pangea. So far, over 26 extinct genera containing over 170 species of these beetles have been described. Three extant genera have been assigned to this family: Omma, Tetraphalerus and Beutelius. The family is considered to be a subfamily of Cupedidae by some authors, but have been found to be more closely related to Micromalthidae in molecular phylogenies. A close relationship with Micromalthidae is supported by several morphological characters, including those of the mandibles and male genitalia. Due to their rarity, their ecology is obscure, it is likely that their larvae feed on deadwood.

David R. Maddison · CC BY 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- CupedoideaPBDB
- Motility
- actively mobilePBDB, for Arthropoda
- Environment
- terrestrialPBDB, for Insecta
Time the record covers
At least 213 million years on record.
The record covers at least 237 Mya to 23.04 Mya. No occurrence read is dated outside 241.464 Mya to 15.98 Mya.
172 occurrences of Ommatidae on record, most of them in the Early Cretaceous. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
172 occurrences- Middle Triassic3%
- Late Triassic2%
- Early Jurassic6%
- Middle Jurassic5%
- Late Jurassic16%
- Early Cretaceous46%
- Late Cretaceous20%
- Eoceneunder 1%
- Mioceneunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
38 formations
- Dzun-Bain Formation · Bon Tsaagan Group22
- Yixian Formation · Jehol Group16
- Karabastau Formation12
- Weald Clay Formation · Wealden Group8
- Solnhofen Formation8
- Daohugou Formation7
Loading coastlines…
The named rock units Ommatidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 14 further formations hold fewer occurrences each and are not listed. 42 occurrences have no formation on record, which is common for older literature.
Ommatidae and its relatives
Early CretaceousWhere Ommatidae and its relatives were recorded in the Early Cretaceous.
Found in the same rock
4,098 taxa- Blattodea8 sites
- Notocupes elegans7 sites
- Allophalerus bontsaganensis6 sites
- Omma liassicum6 sites
- Actea sphinx5 sites
- Aeschnidiidae5 sites
- Aeschnogomphus intermedius5 sites
- Ampliceps furcitibia5 sites
- Chresmoda obscura5 sites
- Cymatophlebia longialata5 sites
- Ditomoptera dubia5 sites
- Hexagenites cellulosus5 sites
- Isophlebia aspasia5 sites
- Lithoblatta lithophila5 sites
- Mesonepa primordialis5 sites
- Myrmicium schroeteri5 sites
- Odonata5 sites
- Omma longicolle5 sites
- Oxycorynoides (Oxycorynoides) similis5 sites
- Protoscelis jurassica5 sites
- Pseudohydrophilus avitus5 sites
- Pycnophlebia speciosa5 sites
- Scarabaeides deperditus5 sites
- Tarsophlebia eximia5 sites
The Paleobiology Database records 95 sites for Ommatidae, and this reads all of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
62 references- New beetles of the family Cupedidae from the Mesozoic of Mongoloa. Ommatini, Mesocupedini, Priacmini22
- New Mesozoic cupedid beetles from Mongolia: Brochocoleini and Notocupedini12
- Brochocolein beetles (Insecta: Coleoptera) from the Lower Cretaceous of northeast China and southern England8
- Ommatin beetles (Insecta: Coleoptera) from the Lower Cretaceous of northeast China and southern England7
- Taxonomic review of fossil coleopterous families (Insecta, Coleoptera). Suborder Archostemata: Superfamilies Coleopseoidea and Cupedoidea6
- Istoricheskoe Razvitie Zhestkokrylykh-Arkhostemat [Historical Development of the Archostomate Beetles]6
- Noviye zhuki semeistva Cupedidae iz Yurskikh otlozheniy Karatau6
- Observations on the beetle family Cupedidae, with descriptions of two new fossil forms and a key to the recent genera6
- Zhuki-Arkhostematy Yury Karatau (Coleoptera, Archostemata) [Jurassic Archostematid beetles from Karatau (Coleoptera, Archostemata)]5
- Cretaceous beetles of the Jinju Formation (Coleoptera): Archostemata4
- New fossil species of ommatids (Coleoptera: Archostemata) from the middle Mesozoic of China illuminating the phylogeny of Ommatidae4
- New ommatids of Ommatinae (Coleoptera: Archostemata: Ommatidae) from the Yixian Formation of Liaoning, China4
Each of the 172 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 50 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Ommatidae
- Source
- Wikipedia/Commons
- Credit
- David R. Maddison
- Licence
- CC BY 3.0
- Commercial use
- the licence allows it
No 3D scan of Ommatidae was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Ommatidae”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itCupedoidea through timeFormations of the Early CretaceousThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Sharp and Muir 1912