Holonematidae
Holonematidae is an extinct family of relatively large arthrodire placoderms from the Early to Late Devonian. Almost all fossil specimens are of armor fragments, though, all have distinctive ornamentation, often of unique arrangements and patterns of tubercles, that are diagnostic of the family. The trunkshield is very elongated, giving the armor an overall "barrel" like appearance.

Apokryltaros · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- PhlyctaeniiformesPBDB
- Diet
- carnivorePBDB, for Arthrodira
- Life habit
- nektonicPBDB, for Arthrodira
- Motility
- actively mobilePBDB, for Arthrodira
Time the record covers
At least 21 million years on record.
The record covers at least 393.47 Mya to 372.15 Mya. No occurrence read is dated outside 419.62 Mya to 358.86 Mya.
34 occurrences of Holonematidae on record, most of them in the Devonian. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
34 occurrences- Devonian100%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
22 formations
- Chemung Formation2
- Narva Formation2
- Aruküla Formation2
- Rockport Quarry Limestone Formation2
- Odenspiel Formation2
- Catskill Formation1
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The named rock units Holonematidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 7 occurrences have no formation on record, which is common for older literature.
Holonematidae and its relatives
DevonianWhere Holonematidae and its relatives were recorded in the Devonian.
Found in the same rock
224 taxa- Holonema11 sites
- Glyptolepis10 sites
- Onychodus7 sites
- Bothriolepis5 sites
- Byssacanthus dilatatus5 sites
- Homostius latus5 sites
- Orvikuina5 sites
- Actinolepis tuberculata4 sites
- Asterolepis estonica4 sites
- Cheirolepis4 sites
- Elasmobranchii4 sites
- Holonema rugosa4 sites
- Asterolepis3 sites
- Cephalaspidida3 sites
- Cheiracanthus brevicostatus3 sites
- Chondrichthyes3 sites
- Coccosteidae3 sites
- Dickosteus3 sites
- Dipterus3 sites
- Ganosteus3 sites
- Ganosteus artus3 sites
- Gyroptychius pauli3 sites
- Heterosteus ingens3 sites
- Holonema obrutshevi3 sites
The Paleobiology Database records 28 sites for Holonematidae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
18 references- End-Devonian extinction and a bottleneck in the early evolution of modern jawed vertebrates5
- The Middle Devonian fishes of the Baltic States (Estonia, Latvia) and Belarus5
- An examination of the Devonian fishes of Michigan4
- On some new and little known Paleozoic vertebrates3
- A new species of the Devonian actinopterygian fish Moythomasia from Belarus2
- Early Devonian arthrodire remains (Placodermi, ?Holonematidae) from the Burrinjuck area, New South Wales, Australia2
- The head shield of Tiaraspis subtilis (Gross) [Pisces, Arthrodira]2
- A new antiarch placoderm from the Emsian (Early Devonian) of Wuding, Yunnan, China1
- Devonian vertebrates from the Canning and Carnarvon Basins with an overview of Paleozoic vertebrates of Western Australia1
- Early Frasnian acanthodians from central Iran1
- Mid- and Late Devonian fossil fishes sites of England and Wales1
- Die arthrodiren Tiaraspis subtilis (Gross) und Dichotiaraspis n. g. (Unterdevon, Emsium)1
Each of the 34 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 6 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Holonematidae
- Source
- Wikipedia/Commons
- Credit
- Apokryltaros
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
No 3D scan of Holonematidae was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itPhlyctaeniiformes through timeFormations of the DevonianThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Obruchev 1932