Enteropneusta
The acorn worms or Enteropneusta are a hemichordate class of invertebrates. The closest non-hemichordate relatives of the Enteropneusta are the echinoderms. There are 111 known species of acorn worm in the world, the main species for research being Saccoglossus kowalevskii. Two families—Harrimaniidae and Ptychoderidae—separated at least 370 million years ago.

Spengel, Johann Wilhelm · Public Domain · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- HemichordataPBDB
Time the record covers
At least 310 million years on record.
The record covers at least 509.7 Mya to 199.5 Mya. No occurrence read is dated outside 516 Mya to 192.9 Mya.
46 occurrences of Enteropneusta on record, most of them in the Cambrian. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
46 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
15 formations
- Burgess Shale Formation · Chancellor Group22
- Kinzers Formation5
- Wheeler Shale Formation3
- Carbondale Formation · Kewanee Group3
- Stephen Formation2
- Balang Formation2
Loading coastlines…
The named rock units Enteropneusta has been recovered from, largest first. Drawn where the rock is now, not where it formed.
Enteropneusta and its relatives
CambrianWhere Enteropneusta and its relatives were recorded in the Cambrian.
Found in the same rock
539 taxa- Canadaspis perfecta21 sites
- Liangshanella21 sites
- Selkirkia columbia21 sites
- Sidneyia21 sites
- Wiwaxia corrugata21 sites
- Diraphora20 sites
- Ehmaniella burgessensis20 sites
- Hazelia20 sites
- Marrella splendens20 sites
- Naraoia compacta20 sites
- Oesia20 sites
- Ottoia prolifica20 sites
- Haplophrentis carinatus19 sites
- Lingulella19 sites
- Micromitra19 sites
- Morania19 sites
- Pentagnostus praecurrens19 sites
- Yohoia19 sites
- Burgessia18 sites
- Scenella amii18 sites
- Trilobita18 sites
- Waptia fieldensis18 sites
- Laggania17 sites
- Molaria17 sites
The Paleobiology Database records 39 sites for Enteropneusta, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
18 references- Paleoecology of the Greater Phyllopod Bed community, Burgess Shale20
- Lower Cambrian Olenellus zone of the Appalachians4
- The diverse radiodont fauna from the Marjum Formation of Utah, USA (Cambrian: Drumian)3
- Saccoglossus testa from the Mazon Creek fauna (Pennsylvanian of Illinois) and the evolution of acorn worms (Enteropneusta: Hemichordata)3
- A Cambrian soft-bodied biota after the first Phanerozoic mass extinction2
- A new phyllopod bed-like assemblage from the Burgess Shale of the Canadian Rockies2
- Tubicolous enteropneusts from the Cambrian period1
- A new, Middle Cambrian, Burgess Shale-type biota, Bolaspidella zone, Chancellor Basin, southeastern British Columbia1
- Palaeontology and depositional environments of ancient brine seeps in the Middle Cambrian Burgess Shale at The Monarch, British Columbia, Canada1
- An unmineralized alga from the Lower Cambrian of California, USA1
- A new genus of Bradoriidae (Crustacea) from the Cambrian of Northern Eurasia1
- New fossil worms from the Lower Cambrian of the Kinzers Formation, Pennsylvania, with some comments on Burgess Shale-type preservation1
Each of the 46 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 6 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Enteropneusta
- Source
- Wikipedia/Commons
- Credit
- Spengel, Johann Wilhelm
- Licence
- Public Domain
- Commercial use
- the licence allows it
No 3D scan was found. Smithsonian Open Access, MorphoSource and Sketchfab were all searched.
The description on this site
- Source
- Wikipedia, “Acorn worm”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itHemichordata through timeFormations of the CambrianThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0.