Dwardius
Dwardius is an extinct genus of cardabiodontid sharks which existed during the Cretaceous period in what is now Australia, England, France, and India. It was described by Mikael Siverson in 1999, as a new genus for the species Cretalamna woodwardi, which had been described by J. Hermann in 1977. Another species, D. siversoni, was described from the middle Albian of northeastern France by V.I. Zhelezko in 2000; the species epithet honours the author of the genus. A new species, D. sudindicus, was described by Charlie J. Underwood, Anjali Goswami, G.V.R. Prasad, Omkar Verma, and John J. Flynn in 2011, from the Cretaceous Karai Formation of India.

Natural History Museum UK · CC0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- CardabiodontidaePBDB
- Diet
- carnivorePBDB, for Lamniformes
- Life habit
- nektonicPBDB, for Lamniformes
- Motility
- actively mobilePBDB, for Lamniformes
Time the record covers
At least 33 million years on record.
The record covers at least 119.57 Mya to 85.7 Mya. No occurrence read is dated outside 121.4 Mya to 83.6 Mya.
20 occurrences of Dwardius on record, most of them in the Late Cretaceous. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
20 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
4 formations
- Hinoshima Formation · Himenoura Group2
- Karai Formation · Uttattur Group1
- Habur Formation1
- Mancos Shale Formation1
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The named rock units Dwardius has been recovered from, largest first. Drawn where the rock is now, not where it formed. 15 occurrences have no formation on record, which is common for older literature.
Dwardius and its relatives
Late CretaceousWhere Dwardius and its relatives were recorded in the Late Cretaceous.
Found in the same rock
150 taxa- Squalicorax falcatus9 sites
- Archaeolamna kopingensis8 sites
- Cretalamna appendiculata7 sites
- Heterodontus canaliculatus7 sites
- Paranomotodon angustidens7 sites
- Pseudoscapanorhynchus compressidens7 sites
- Squatina (Cretascyllium) cranei7 sites
- Synechodus dubrisiensis7 sites
- Heterodontus6 sites
- Squatina muelleri6 sites
- Protolamna5 sites
- Ptychodus mammillaris5 sites
- Scapanorhynchus minimus5 sites
- Anomotodon principalis4 sites
- Cretodus semiplicatus4 sites
- Cretoxyrhina mantelli4 sites
- Eostriatolamia subulata4 sites
- Leptostyrax macrorhiza4 sites
- Paraisurus macrorhizus4 sites
- Paraorthacodus recurvus4 sites
- Ptychodus decurrens4 sites
- Squalicorax4 sites
- Squatirhina thiesi4 sites
- Synechodus tenuis4 sites
The Paleobiology Database records 20 sites for Dwardius, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
11 references- Les Elasmobranches de l'Albien inferieur et moyen (Cretace inferieur) de la Marne et de la Haute-Marne (France)4
- Sharks (Elasmobranchii: Euselachii) from the Late Cretaceous of France and the UK3
- Features and paleoecological significance of the shark fauna from the Upper Cretaceous Hinoshima Formation, Himenoura Group, southwest Japan2
- Mid-Cretaceous Cretoxyrhina (Elasmobranchii) from Mangyshlak, Kazakhstan and Texas, USA2
- Sélaciens nouveaux du Crétacé supérieur du Sud-Ouest de la France. Quelques apports à la systématique des élasmobranches2
- Les sélaciens des terrains néocrétacés and paléocènes de Belgique and des contrées limitrophes. Eléments d'une biostratigraphie intercontinentale2
- First Early Cretaceous sharks from India1
- Late late Albian (Early Cretaceous) shark teeth from Annopol, Poland1
- Marine vertebrates from the ‘Middle’ Cretaceous (Early Cenomanian) of South India1
- The upper Albian and lower Cenomanian succession at Kolbay, eastern Mangyshlak (southwest Kazakhstan)1
- Selachians from the Greenhorn Cyclothem ("Middle" Cretaceous: Cenomanian-Turonian), Black Mesa, Arizona, and the paleogeographic distribution of Late Cretaceous selachians1
Each of the 20 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Dwardius
- Source
- Wikipedia/Commons
- Credit
- Natural History Museum UK
- Licence
- CC0
- Commercial use
- the licence allows it
No 3D scan of Dwardius was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Dwardius”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itCardabiodontidae through timeFormations of the Late CretaceousThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Siverson 1999