Borophagus pugnator
Borophagus pugnator is an extinct species of the genus Borophagus of the subfamily Borophaginae, a group of canids endemic to North America from the early Miocene epoch through the late Miocene epoch from 10.3 to 9.4 Ma. Borophagus pugnator existed for approximately 0.9 million years.

Geekgecko · CC BY 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- BorophagusPBDB
- Diet
- carnivorePBDB, for Borophagus
- Life habit
- ground dwellingPBDB, for Canidae
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Eutheria
Time the record covers
At most 7.8 million years on record.
Every occurrence read is dated to somewhere between 12.5 Mya and 4.7 Mya, and nothing in the dating separates them, so that window is what the record shows rather than a span.
18 occurrences of Borophagus pugnator on record, most of them in the Miocene. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
18 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
4 formations
- Peace River Formation · Hawthorn Group5
- Ogallala Formation2
- Alachua Formation1
- Poison Creek Formation1
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The named rock units Borophagus pugnator has been recovered from, largest first. Drawn where the rock is now, not where it formed. 9 occurrences have no formation on record, which is common for older literature.
Borophagus pugnator and its relatives
MioceneWhere Borophagus pugnator and its relatives were recorded in the Miocene.
Found in the same rock
182 taxa- Epicyon haydeni7 sites
- Neohipparion eurystyle5 sites
- Aphelops mutilus4 sites
- Protohippus gidleyi4 sites
- Amphimachairodus coloradensis3 sites
- Camelidae3 sites
- Cormohipparion ingenuum3 sites
- Enhydritherium terraenovae3 sites
- Floridameryx floridanus3 sites
- Hexameryx simpsoni3 sites
- Nannippus aztecus3 sites
- Serridentinus floridanus3 sites
- Tayassuinae3 sites
- Teleoceras3 sites
- Teleoceras hicksi3 sites
- Aepycamelus major2 sites
- Aphelops malacorhinus2 sites
- Borophagus hilli2 sites
- Calippus2 sites
- Cormohipparion emsliei2 sites
- Cormohipparion plicatile2 sites
- Corystosiren varguezi2 sites
- Dinohippus interpolatus2 sites
- Dinohippus mexicanus2 sites
The Paleobiology Database records 18 sites for Borophagus pugnator, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
5 references- Phylogenetic systematics of the Borophaginae (Carnivora: Canidae)13
- Synonymies and reidentifications of North American fossil mammals2
- Stop 15: Early Hemphillian faunas of the Texas and Oklahoma panhandles1
- Stratigraphy of the Ogallala Formation (Neogene) of northern Kansas1
- Tertiary Land Mammals of Florida1
Each of the 18 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. Each entry links to the Paleobiology Database's own record for the work.
Media and terms
Media content1
Picture of Borophagus pugnator
- Source
- Wikipedia/Commons
- Credit
- Geekgecko
- Licence
- CC BY 4.0
- Commercial use
- the licence allows it
No 3D scan of Borophagus pugnator was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itBorophagus through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Cook 1922)